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Cnidaria

Cnidaria (〔''Dictionary.com Unabridged.'' Random House, Inc. (Cnidaria ). Retrieved May 15, 2013.〕) is a phylum containing over 10,000 species of animals found exclusively in aquatic (freshwater and marine) environments: they are predominantly marine species. Their distinguishing feature is cnidocytes, specialized cells that they use mainly for capturing prey. Their bodies consist of mesoglea, a non-living jelly-like substance, sandwiched between two layers of epithelium that are mostly one cell thick. They have two basic body forms: swimming medusae and sessile polyps, both of which are radially symmetrical with mouths surrounded by tentacles that bear cnidocytes. Both forms have a single orifice and body cavity that are used for digestion and respiration. Many cnidarian species produce colonies that are single organisms composed of medusa-like or polyp-like zooids (hence they are trimorphic), or both. Cnidarians' activities are coordinated by a decentralized nerve net and simple receptors. Several free-swimming Cubozoa and Scyphozoa possess balance-sensing statocysts, and some have simple eyes. Not all cnidarians reproduce sexually, with many species having complex life cycles of asexual polyp stages and sexual medusae. Some, however, omit either the polyp or the medusa stage.
Cnidarians were formerly grouped with Ctenophores in the phylum Coelenterata, but increasing awareness of their differences caused them to be placed in separate phyla. Cnidarians are classified into four main groups: the almost wholly sessile Anthozoa (sea anemones, corals, sea pens); swimming Scyphozoa (jellyfish); Cubozoa (box jellies); and Hydrozoa, a diverse group that includes all the freshwater cnidarians as well as many marine forms, and has both sessile members, such as ''Hydra'', and colonial swimmers, such as the Portuguese Man o' War. Staurozoa have recently been recognised as a class in their own right rather than a sub-group of Scyphozoa, and there is debate about whether Myxozoa and Polypodiozoa are cnidarians or closer to bilaterians (more complex animals).
Most cnidarians prey on organisms ranging in size from plankton to animals several times larger than themselves, but many obtain much of their nutrition from endosymbiotic algae, and a few are parasites. Many are preyed on by other animals including starfish, sea slugs, fish and turtles. Many scleractinian corals—which form the structural foundation for coral reefs—possess polyps that are filled with endosymbiotic algae. While reef-forming corals are almost entirely restricted to warm and shallow marine waters, other cnidarians can be found at great depths, in polar regions, and in freshwater.
Fossil cnidarians have been found in rocks formed about , and other fossils show that corals may have been present shortly before and diversified a few million years later. Fossils of cnidarians that do not build mineralized structures are very rare. Scientists currently think that cnidarians, ctenophores and bilaterians are more closely related to calcareous sponges than these are to other sponges, and that anthozoans are the evolutionary "aunts" or "sisters" of other cnidarians, and the most closely related to bilaterians.
==Distinguishing features==

Cnidarians form an animal phylum that are more complex than sponges, about as complex as ctenophores (comb jellies), and less complex than bilaterians, which include almost all other animals. However, both cnidarians and ctenophores are more complex than sponges as they have: cells bound by inter-cell connections and carpet-like basement membranes; muscles; nervous systems; and some have sensory organs. Cnidarians are distinguished from all other animals by having cnidocytes that fire like harpoons and are used mainly to capture prey. In some species, cnidocytes can also be used as anchors.
Like sponges and ctenophores, cnidarians have two main layers of cells that sandwich a middle layer of jelly-like material, which is called the mesoglea in cnidarians; more complex animals have three main cell layers and no intermediate jelly-like layer. Hence, cnidarians and ctenophores have traditionally been labelled diploblastic, along with sponges.〔〔 However, both cnidarians and ctenophores have a type of muscle that, in more complex animals, arises from the middle cell layer. As a result, some recent text books classify ctenophores as triploblastic,〔 and it has been suggested that cnidarians evolved from triploblastic ancestors.〔

抄文引用元・出典: フリー百科事典『 ウィキペディア(Wikipedia)
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