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Areole : ウィキペディア英語版
Areole

In botany, areoles are small light- to dark-colored bumps on cacti out of which grow clusters of spines. Areoles are important diagnostic features of cacti, and identify them as a family distinct from other succulent plants.〔Anderson, Edward F., The Cactus Family, Pub: Timber Press 2001 ISBN 978-0-88192-498-5〕 The spines are not easily detachable, but on certain cacti, members of the subfamily Opuntioideae, smaller, detachable bristles, glochids, also grow out of the areoles and afford additional protection.
Areoles represent highly specialized branches on cacti. Apparently, they evolved as abortive branch buds while their spines evolved as vestigial leaves. In branched cacti, such as Opuntioidiae and the saguaro, new branches grow from areoles, because that is where the buds are. The development of the areole seems to have been an important element in the adaptation of cacti to niches in desert ecology.
Some of the Opuntioideae have spines, as well as glochids, on their areoles; some have only glochids. Structurally, the glochids seem to be bristles rather than evolved leaves. They are detachable and resemble small, sharp splinters. Unlike the spines, glochids generally are barbed and are very difficult to remove from the skin.
==Evolution==
The cactus family, the Cactaceae, evolved 30–40 million years ago in the Americas, originally completely separately from Africa, Europe, and Asia, although, probably within the last few million years, some species of ''Rhipsalis'' appear to have been carried to parts of Asia and Africa, most likely by birds.
The areole was one of the unique features that cacti developed, possibly in adaptation to the ambient climate in the area in which they emerged. Accordingly, the areoles distinguish the family Cactaceae from other succulent plants both in the New World and the Old World.

抄文引用元・出典: フリー百科事典『 ウィキペディア(Wikipedia)
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