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Boraginoideae
Boraginoideae is a subfamily of the flowering plant family Boraginaceae, as that family is defined in the APG III system of classification for flowering plants. The Angiosperm Phylogeny Group has not actually specified subfamilies within Boraginaceae. Some taxonomists place the genera ''Codon'' and ''Wellstedia'' in Boraginoideae. Others place one or both of these in separate, monogeneric subfamilies. ''Codon'' was long regarded as an odd member of Hydrophylloideae, but in 1998, a molecular phylogenetic study showed that it is closer to Boraginoideae.〔Diane M. Ferguson. 1998. "Phylogenetic Analysis and Relationships in Hydrophyllaceae Based on ndhF Sequence Data". ''Systematic Botany'' 23(3):253-268.〕
Many of the botanists who work with Boraginaceae do not follow the APG III system.〔Maximilian Weigend, Federico Luebert, Marc Gottschling, Thomas L.P. Couvreur, Hartmut H. Hilger and James S. Miller. 2014. "From capsules to nutlets — phylogenetic relationships in the Boraginales". ''Cladistics'' 30(5):508-518. .〕 Instead, they recognize five to eight families in the order Boraginales.〔Peter F. Stevens. (2001 onwards). "Boraginaceae". At: Angiosperm Phylogeny Website. At: Missouri Botanical Garden Website. (see ''External links'' below)〕
Comparisons of DNA sequences by cladistic methods have strongly supported the division of Boraginoideae into four tribes: Echiochileae, Boragineae, Lithospermeae, and Cynoglosseae.〔James I. Cohen. 2014. "A phylogenetic analysis of morphological and molecular characters of Boraginaceae: evolutionary relationships, taxonomy, and patterns of character evolution". ''Cladistics'' 30(2):139-169. 〕
== Genera ==
The following list of genera consists of ''Codon'' plus the genera listed for Boraginoideae at the Germplasm Resources Information Network
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