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Epsilonproteobacteria are a class of Proteobacteria. All species of this class are, like all Proteobacteria, Gram-negative. The Epsilonproteobacteria consist of few known genera, mainly the curved to spirilloid ''Wolinella'' spp., ''Helicobacter'' spp., and ''Campylobacter'' spp. Most of the known species inhabit the digestive tracts of animals and serve as symbionts (''Wolinella'' spp. in cattle) or pathogens (''Helicobacter'' spp. in the stomach, ''Campylobacter'' spp. in the duodenum). Numerous environmental sequences and isolates of Epsilonproteobacteria have also been recovered from hydrothermal vents and cold seep habitats. Examples of isolates include ''Sulfurimonas autotrophica'', ''Sulfurimonas paralvinellae'',''Sulfurovum lithotrophicum'' and ''Nautilia profundicola''. A member of the class Epsilonproteobacteria occurs as an endosymbiont in the large gills of the deepwater sea snail ''Alviniconcha hessleri''.〔Yohey Suzuki, Takenori Sasaki, Masae Suzuki, Yuichi Nogi, Tetsuya Miwa, Ken Takai, Kenneth H. Nealson & Koki Horikoshi (September 2005) (Novel Chemoautotrophic Endosymbiosis between a Member of the ''Epsilonproteobacteria'' and the Hydrothermal-Vent Gastropod ''Alviniconcha'' aff. ''hessleri'' (''Gastropoda'': ''Provannidae'') from the Indian Ocean ). ''Applied and Environmental Microbiology'', 71(9): 5440-5450.〕 The Epsilonproteobacteria found at deep-sea hydrothermal vents characteristically exhibit chemolithotrophy, meeting their energy needs by oxidixing reduced sulfur, formate, or hydrogen coupled to the reduction of nitrate or oxygen. Autotrophic Epsilonproteobacteria use the reverse Krebs cycle to fix carbon dioxide into biomass, a pathway originally thought to be of little environmental significance. The oxygen sensitivity of this pathway is consistent with their microaerophilic or anaerobic niche in these environments, and their likely evolution in the Mesoproterozoic oceans, which are thought to have been sulfidic with low levels of oxygen available from cyanobacterial photosynthesis. ==Phylogeny== The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) and National Center for Biotechnology Information (NCBI) and the phylogeny is based on 16S rRNA-based LTP release 106 by 'The All-Species Living Tree' Project 〔'The All-Species Living Tree' Project.(【引用サイトリンク】 title=16S rRNA-based LTP release 106 (full tree) )〕 }} |2= |4= }} }} }} }} }} }} Notes: * Prokaryotes where no pure (axenic) asses are isolated or available, i.e. not cultivated or can not be sustained in culture for more than a few serial passages 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Epsilonproteobacteria」の詳細全文を読む スポンサード リンク
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