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Haptopoda
Haptopoda is an extinct arachnid order known exclusively from only nine specimens from the Upper Carboniferous of Coseley, Staffordshire, United Kingdom. Haptopoda is monotypic, represented only by the species ''Plesiosiro madeleyi'' described by Reginald Innes Pocock in his important 1911 monograph on British Carboniferous arachnids. The original locality from which these fossils originate is no longer available thus it is unclear whether any further examples will be found.
The original fossils have been redescribed in detail by Alexander Petrunkevitch in 1949 and Dunlop in 1999. A supposed example from the Coal Measures of Lancashire is a misidentification.
==Relationships==
Relationships with other arachnids are obscure. ''Plesiosiro'' means "close to ''Siro''", which is a genus of cyphophthalmid (Cyphophthalmi); the most primitive group of the living harvestmen (Opiliones). These harvestmen do, in some ways, resemble the reconstructed body plan of the haptopods.
Revisions have confirmed that Haptopoda should be treated as a separate and independent order. The most recent study tentatively recognised a group named Schizotarsata Shultz, 2007 comprising Haptopoda, Amblypygi (whip spiders), Uropygi (whip scorpions) and Schizomida (schizomids). All share the character of a subdivided tarsus (or foot) which gives the group its name. Some cladistic analyses recover the phalangiotarbids as sister group to the Opiliones, albeit with low support.
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