|
''Horneophyton'', a member of the Horneophytopsida, is an extinct early plant which may form a "missing link" between the hornworts and the Rhyniopsida. It is among the most abundant organisms found in the Rhynie chert, a Devonian Lagerstätte in Scotland. ==Description== The sporophyte had bare stems (axes) up to 20 cm high and about 2 mm in diameter with an undivided cortex; stomata were present but rare. There was a thin central strand of conducting tissue, but this was not reinforced with spiral and reticulate thickenings (and thus does not constitute true vascular tissue). Early stages of development of the sporophytes of ''Horneophyton'' (as of hornworts) may have been dependent on their parent gametophytes for nutrition, but mature specimens have expanded, corm-like bases to their stems, up to 6 mm in diameter, that bore rhizoids and appear to be anchored in soil, suggesting a capacity for independent existence after the gametophyte had degenerated. The sporangium (spore-forming organ) is unique among both living and fossil plants, consisting as it does of branched lobes at the apex of some of the branches of the stem. Each lobe contains a central collumella, analogous to the sporangia of hornworts; however, the sporangia of hornworts do not dichotomise.〔, pp. 237–8〕 The number of lobes possessed by a sporangium varied; at least three orders of dichotomous branching have been found, resulting in more than four lobes. The sporangia were much less regular than shown in most reconstructions (including that opposite), and they had 'bumps' or emergences on them. Spores were released through a slit at the top of each lobe. The sporangia of ''Horneophyton'' contained trilete meiospores, the surfaces of which were decorated with short conical protuberances.〔 The female gametophyte of the plant has been recognised and described as the form taxon ''Langiophyton mackiei''. It grew to a height of around 6 cm, and was free living. It may have been dioicous, i.e. produced male and female gametes on separate plants. ''Horneophyton'' grew on sandy, organic-rich soil in damp to wet locations. They usually grew as isolated individuals. 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Horneophyton」の詳細全文を読む スポンサード リンク
|