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Hulkepholis
''Hulkepholis'' is an extinct genus of goniopholidid mesoeucrocodylian from the Early Cretaceous of southern England and eastern Spain. It contains two species, the type species, ''Hulkepholis willetti'', and also ''H. plotos''. ''Hulkepholis'' is most closely related to both species of ''Anteophthalmosuchus'' and to the unnamed "Dollo’s goniopholidid".
==Discovery==
''H. willetti'' is known from a single holotype specimen, BMNHB 001876 from the Wealden Group of the Isle of Wight that includes a well-preserved nearly complete skull.〔 It was collected by Edgar W. Willett at Cuckfield, West Sussex, from the Valanginian-aged Grinstead Clay Formation, of the Hastings Group, Wealden Supergroup. Willett showed the specimen to the Geological Society of London in or around 1877 or 1878, and it was then Hulke (1878) described it as a specimen of ''Goniopholis crassidens'', an identification derived from a similar tooth form. The specimen was considered lost among the crocodyliform research community but was actually safely accessioned at the Booth Museum in Brighton. It was identified as "the long-lost crocodilian of Mr Willett" by Steve (1998), "Willett’s specimen" by Salisbury ''et al.'' (1999) and "Hulke’s specimen" by Andrande ''et al.'' (2011). It was assigned to a new species of ''Goniopholis'', ''Goniopholis willetti'', by Salisbury and Naish in 2011 honoring its collector.〔 A broad phylogenetic analysis of crocodyliforms published the same year by Andrande ''et al.'' (2011), found the specimen to be the sister taxon of the clade formed by ''Anteophthalmosuchus hooleyi'' and the unnamed "Dollo’s goniopholidid".〔 Thus it was reassigned to its own genus by Buscalioni ''et al.'' (2013), creating the ''combinatio nova'', ''Hulkepholis willetti''. The generic name honors John Whitaker Hulke who described the specimen as belonging to ''Goniopholis''.〔
A second species, ''Hulkepholis plotos'', was first described and named by A.D. Buscalioni, L. Alcalá, E. Espílez and L. Mampel in 2013. The specific name is derived from Greek mythology ''πλοτός'', ''plotos'', the drifter. It is known solely from the holotype AR-1/56, a partial skeleton which consists of AR-1-2045, a nearly complete but crushed skull; AR-1-2048, 4859, 4860, three vertebrae; AR-1-2046, a rib; AR-1-2048, a metapodial; and AR-1-2049, 4861, 4862, three osteoderms. It was collected from the early Albian-aged Escucha Formation, at Santa Maria Mine located in the municipality of Ariño, Teruel Province, of Aragon, along with the closely related ''Anteophthalmosuchus escuchae''.〔
抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』
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