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Lepidobotryaceae is a flowering plant family in the order Celastrales.〔"Lepidobotryaceae" In: Peter F. Stevens (2001 onwards). Angiosperm Phylogeny Website. In: Missouri Botanical Garden Website. (see External links below).〕 It contains only two species, ''Lepidobotrys staudtii'' and ''Ruptiliocarpon caracolito''.〔Klaus Kubitzky. "Lepidobotryaceae" In: Klaus Kubitzki (ed.). ''The Families and Genera of Vascular Plants'' vol.VI. Springer-Verlag: Berlin,Heidelberg, Germany (2004).〕 == Description == The Lepidobotryaceae are dioecious trees. The leaves are alternate and arranged in two rows along the stems. The blade is elliptic in shape and the margin is entire. The leaves appear simple, but are actually unifoliate. A unifoliate leaf is a type of compound leaf that consists of a single leaflet mounted on the end of a rachis. There is a joint where the leaflet is attached to the rachis.〔Glossary In: Peter F. Stevens (2001 onwards). Angiosperm Phylogeny Website. In: Missouri Botanical Garden Website. (see External links below).〕 In Lepidobotryaceae, this joint bears a single, elongate stipel, and there is a pair of small stipules where the petiole attaches to the stem. After the emergence of the leaf, the stipel and stipules soon fall away. The flowers are produced in small inflorescences opposite the leaves.〔Barry E. Hammel, and Nelson A. Zamora (1993). "''Ruptiliocarpon'' (Lepidobotryaceae): A New Arborescent Genus and Tropical American Link to Africa, with a Reconsideration of the Family". ''Novon'' 3(4):408-417.〕 They are small and greenish with five sepals and five petals. The sepals and petals are similar in size and appearance, free from each other, or very shortly united at the base. In the flower bud, the sepals are arranged quincuncially. This means that two are inside, two are outside, and one of them has one margin exposed and the other covered.〔Benjamin D. Jackson. ''A Glossary of Botanic Terms''. Duckworth: London (1928).〕 The nectary disk is fleshy in ''Lepidobotrys'', but extended into a tube in ''Ruptiliocarpon''.〔 The stamens are in two whorls of five, one whorl opposite the sepals and the other opposite the petals. Those in the outer whorl, opposite the sepals, are longer. The filaments are fused at the base, shortly in ''Lepidobotrys'', but forming an extension of the tubular nectary in ''Ruptiliocarpon''. The pollen is produced in four thecae on each anther. The stigmas are elongate, appearing as false styles, known as stylodia.〔 The ovary is located inside the flower, rather than below. It has two or three locules, with two ovules per locule. The ovules are attached to the partition that separates the locules, near its summit. The fruit is a capsule with one, or rarely, two seeds. The seeds are black and partly covered with an orange aril. In 2000, a DNA analysis of the eudicots based on the rbcL gene showed that the families Lepidobotryaceae, Parnassiaceae, and Celastraceae form a strongly supported clade.〔Vincent Savolainen, Michael F. Fay, Dirk C. Albach, Anders Backlund, Michelle van der Bank, Kenneth M. Cameron, S.A. Johnson, M. Dolores Lledo, Jean-Christophe Pintaud, Martyn P. Powell, Mary Clare Sheahan, Douglas E. Soltis, Pamela S. Soltis, Peter Weston, W. Mark Whitten, Kenneth J. Wurdack and Mark W. Chase (2000). "Phylogeny of the eudicots: a nearly complete familial analysis based on ''rbcL'' gene sequences". ''Kew Bulletin'' 55(2):257-309.〕 The authors of this study recommended that these three families constitute the order Celastrales. This result was strongly supported by later studies.〔Li-Bing Zhang and Mark P. Simmons (2006). "Phylogeny and Delimitation of the Celastrales Inferred from Nuclear and Plastid Genes". ''Systematic Botany'' 31(1):122-137.〕〔Hengchang Wang, Michael J. Moore, Pamela S. Soltis, Charles D. Bell, Samuel F. Brockington, Roolse Alexandre, Charles C. Davis, Maribeth Latvis, Steven R. Manchester, and Douglas E. Soltis (2009). "Rosid radiation and the rapid rise of angiosperm-dominated forests". ''Proceedings of the National Academy of Sciences'' 106(10):3853-3858. 10Mar2009.〕 The families that ''Lepidobotrys'' had usually been placed into, Linaceae and Oxalidaceae, are now placed in the orders Malpighiales and Oxalidales, respectively, which are closely related to Celastrales. The orders Celastrales, Oxalidales, and Malpighiales, along with the unplaced family Huaceae form a group known as the COM clade of the Rosids.〔 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Lepidobotryaceae」の詳細全文を読む スポンサード リンク
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