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The Myrtaceae are a family, the myrtle family, of dicotyledonous plants placed within the order Myrtales. Myrtle, pohutukawa, the bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus all are members of this group. All species are woody, with essential oils, and flower parts in multiples of four or five. One notable character of the family is that the phloem is located on both sides of the xylem, not just outside as in most other plants. The leaves are evergreen, alternate to mostly opposite, simple, and usually with an entire (not toothed) margin. The flowers have a base number of five petals, though in several genera the petals are minute or absent. The stamens are usually very conspicuous, brightly coloured and numerous. == Diversity == Recent estimates suggest the Myrtaceae include over 5650 species,〔Govaerts, R. et al. (12 additional authors). 2008. World Checklist of Myrtaceae. Royal Botanic Gardens, Kew. xv + 455 pp.〕 occurring in some 130-150 genera. The family has a wide distribution in tropical and warm-temperate regions of the world, and are typically common in many of the world's biodiversity hotspots. Genera with capsular fruits such as ''Eucalyptus'', ''Corymbia'', ''Angophora'', ''Leptospermum'', ''Melaleuca'', and ''Metrosideros'' are absent from the Americas, apart from the monotypic genus ''Tepualia'' in Chile and Argentina. Genera with fleshy fruits have their greatest concentrations in eastern Australia and Malesia (the Australasia ecozone) and the Neotropics. ''Eucalyptus'' is a dominant, nearly ubiquitous genus in the more mesic parts of Australia and extends north sporadically to the Philippines. ''Eucalyptus regnans'' is the tallest flowering plant in the world. Other important Australian genera are ''Callistemon'' (bottlebrushes), ''Syzygium'', and ''Melaleuca'' (paperbarks). Species of the genus ''Osbornia'', native to Australasia, are mangroves. ''Eugenia'', ''Myrcia'', and ''Calyptranthes'' are among the larger genera in the neotropics. Historically, the Myrtaceae were divided into two subfamilies. Subfamily Myrtoideae (about 75 genera) was recognized as having fleshy fruits and opposite, entire leaves. Most genera in this subfamily have one of three easily recognized types of embryos. The genera of Myrtoideae can be very difficult to distinguish in the absence of mature fruits. Myrtoideae are found worldwide in subtropical and tropical regions, with centers of diversity in the Neotropics, northeastern Australia, and Malesia. In contrast, subfamily Leptospermoideae (about 80 genera) was recognized as having dry, dehiscent fruits (capsules) and leaves arranged spirally or alternate. The Leptospermoideae are found mostly in Australasia, with a centre of diversity in Australia. Many genera in Western Australia have greatly reduced leaves and flowers typical of more xeric habitats. The division of Myrtaceae into Leptospermoideae and Myrtoideae was challenged by a number of authors, including Johnson and Briggs (1984), who identified 14 tribes or clades within Myrtaceae, and found Myrtoideae to be polyphyletic. Molecular studies by several groups of authors, as of 2008, have confirmed the baccate (fleshy) fruits evolved twice from capsular fruits, and as such, the two-subfamily classification does not accurately portray the phylogenetic (= evolutionary) history of the family. Thus, many workers are now using a recent analysis by Wilson et al. (2001) as a starting point to test further analyses of the family. The genera ''Heteropyxis'' and ''Psiloxylon'' have been separated as separate families by many authors in the past as Heteropyxidaceae and Psiloxylaceae.〔Sytsma, Kenneth J. and Amy Litt. 2002. Tropical disjunctions in and among the Myrtaceae clade (Myrtaceae, Heteropyxidaceae, Psiloxylaceae, Vochysiaceae): Gondwanan vicariance or dispersal? (Abstract). Botany 2002 Conference, University of Wisconsin, Madison, Wisconsin, August 4–7, 2002.〕〔Johnson, L.A.S. and Briggs, B.G. 1984. Myrtales and Myrtaceae – a phylogenetic analysis. ''Annals of the Missouri Botanic Garden'' 71: 700-756.〕 However, Wilson ''et al.''〔 Wilson, Peter G., O'Brien, Marcelle M., Gadek, Paul A., and Quinn, Christopher J. 2001. "Myrtaceae Revisited: A Reassessment of Infrafamilial Groups". ''American Journal of Botany'' 88 (11): 2013–2025. Available (online ) (pdf file).〕 included them in Myrtaceae. These two genera are presently believed to be the earliest arising and surviving lineages of Myrtaceae. The most recent classification recognizes 17 tribes and two subfamilies, Myrtoideae and Psiloxyloideae, based on a phylogenetic analysis of plastid DNA.〔Wilson, P.G., O’Brien, M.M., Heslewood, M.M., Quinn, C.J. 2005. Relationships within Myrtaceae sensu lato based on a matK phylogeny. ''Plant Systematics and Evolution'' 251: 3–19.〕 Many new species are being described annually from throughout the range of Myrtaceae. Likewise, new genera are being described nearly yearly. 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Myrtaceae」の詳細全文を読む スポンサード リンク
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