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List of fossil primates
This is a list of fossil primates—extinct primates for which a fossil record exists. Primates are generally thought to have evolved from a small, unspecialized mammal, which probably fed on insects and fruits. However, the precise source of the primates remains controversial and even their arboreal origin has recently been questioned. As it has been suggested, many other mammal orders are arboreal too, but they have not developed the same characteristics as primates. Nowadays, some well known genera, such as ''Purgatorius'' and ''Plesiadapis'', thought to be the most ancient primates for a long time, are not usually considered as such by recent authors, who tend to include them in the new order Plesiadapiformes, within superorder Euarchontoglires. Some, to avoid confusions, employ the unranked term Euprimates, which excludes Plesiadapiformes. That denomination is not used here.
There is an academic debate on the time the first primates appeared. One of the earliest probable primate fossils is the problematic ''Altiatlasius koulchii'', perhaps an Omomyid, but perhaps a non-Primate Plesiadapiform, which lived in Morocco, during the Paleocene, around 60 Ma.〔 However, other studies, including molecular clock studies, have estimated the origin of the primate branch to have been in the mid-Cretaceous period, around 85 Ma, that is to say, in the time previous to the extinction of dinosaurs and the successful mammal radiation. Nevertheless, there seems to be a consensus about the monophyletic origin of the order, although the evidence is not clear. There are no fossils known that can be directly linked to the living African apes, nor any that could be considered representative of the last common ancestor between them and humans.〔Willoughby, P. Palaeoanthropology and the place of humans in nature. International Journal of Comparative Psychology, 2005, 18, 60-91. Pamela Willougby at the time of authorship is Prof of Anthropology at the University of Alberta〕
The order Primates, established by Linnaeus in 1758, includes humans and their immediate ancestors. However, contrarily to the common opinion, most primates do not have especially large brains. Brain size is a derived character, which only appeared with genus ''Homo'', and was lacking in the first hominid. In fact, hominid encephalization quotient is only 1.5 Ma more recent than that of some dolphin species. The encephalization quotient of some cetaceans is therefore higher than that of most primates, including the nearest relatives of humans, such as ''Australopithecus''.
This list follows partly from Walter Carl Hartwig's 2002 book ''The Fossil Primate Record'' and John G. Fleagle's 2013 book ''Primate Adaptation and Evolution'' (3rd edition). Parentheses around authors' names (and dates) indicates a change in generic name for the fossil, as stated in the International Code of Zoological Nomenclature (ICZN). Since the publication of the book as well as the creation of this article, new fossil taxon have been discovered that has helped improved the taxonomy among primates in general.
== Strepsirrhini ==
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