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Odontogriphus : ウィキペディア英語版 | Odontogriphus
''Odontogriphus'' (literally "toothed riddle") is a genus of soft-bodied animals known from middle Cambrian Lagerstätte. Reaching as much as in length, ''Odontogriphus'' is a flat, oval bilaterian which apparently had a single muscular foot, and a "shell" on its back that was moderately rigid but of a material unsuited to fossilization. Originally it was known from only one specimen, but 189 new finds in the years immediately preceding 2006 made a detailed description possible. () As a result, ''Odontogriphus'' has become prominent in the debate that has gone on since 1990 about the evolutionary origins of molluscs, annelid worms and brachiopods. A group of scientists think that ''Odontogriphus''’s feeding apparatus, which is "nearly identical" to Wiwaxia’s, is an early version of the molluscan radula, a chitinous "tongue" that bears multiple rows of rasping teeth. Hence they classify ''Odontogriphus'' and ''Wiwaxia'' as close to the ancestors of the first true molluscs. One scientist has presented a different analysis, arguing since 1990 that ''Wiwaxia'' is not closely related to molluscs but is much more like a polychaete worm. He argues that the supposed "radula" is nothing of the sort; he classifies ''Odontogriphus'' as a basal lophotrochozoan, in other words close to the last common ancestor of molluscs, annelid worms and brachiopods. ==History of discovery==
Charles Doolittle Walcott found one specimen during one of his field trips to the Burgess Shale between 1910 and 1917. In the 1970s Simon Conway Morris re-examined the specimen and tentatively concluded that it was a swimming lophophorate, in other words related to the ancestors of molluscs, annelid worms and brachiopods. In 2006 Caron, Scheltema ''et al.'' published a new analysis based on 189 recently collected specimens, all from the Burgess Shale.〔 A full pre-publication draft, free but without pictures, may be available at (【引用サイトリンク】format=PDF )〕
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