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Pachypteris
''Pachypteris'' (Brongn.) T.M.Harris. is a Mesozoic pteridosperm leaf fossil probably belonging to the seed fern Order Corystospermales. ''Pachypteris'' is represented by hypostomatic, bipinnate or unipinnate leaves, with alethopteridian venation (midvein and secondary veins divided once or twice before reaching the pinnule margin), pinnules with entire margins and rounded apices. The stomata are haplocheilic, monocyclic or dicyclic, usually depressed, with the guard cells occurring in the lowermost part of the stoma. The affinities of ''Pachypteris'' lay with ''Cycadopteris'', ''Komlopteris'', ''Dicroidium'' (a typical Corystospermalean foliage) and ''Ptilozamites''. It includes the former denomination ''Thinnfeldia'' Ettingshausen 1852, a junior synonym of ''Pachypteris'', as Doludenko (1971) 〔Doludenko, M.P., 1971. Thinnfeldia - mladsii sinonim ''Pachypteris''. Paleontological Journal, 2: 99-104.〕 showed. The genus was detailed by Harris (1964),〔Harris, T.M., 1964. The Yorkshire Jurassic Flora. Part 2: Caytoniales, Cycadales and Pteridospermales, 2. British Museum (Natural History), London, 1-192 pp.〕 Doludenko (1974),〔Doludenko, M.P., 1974. On the relation of the genera ''Pachyperis'', ''Thinnfeldia'' and ''Cycadopteris'', Symposium on Morphological and Stratigraphical Paleobotany. Birbal Sahni Institute of Paleobotany, Lucknow, pp. 8-16.〕 Schweitzer and Kirchner (1998),〔Schweitzer, H.-J. and Kirchner, M., 1998. Die Rhato-Jurassischen Floren des Iran und Afghanistans: 11. Pteridospermophyta und Cycadophyta. I. Cycadales. Palaeontographica Abt. B, 248: 1-85.〕 Popa (2000),〔Popa, M.E., 2000. Aspects of Romanian Early Jurassic palaeobotany and palynology. Part II. A new species of ''Pachypteris'' from Cristian. Review of Palaeobotany and Palynology, 111: 31-47.〕 and Gordenko (2007).〔 The genus ''Komlopteris'', a segregate from ''Pachypteris'', was defined by Barbacka (1994).〔Barbacka, M., 1994. ''Komlopteris'' Barbacka, gen. nov., a segregate from ''Pachypteris'' Brongniart. Review of Palaeobotany and Palynology(83): 339-349.〕 ''Pachypteris'' includes about 20 species ranging from late Triassic to Lower Cretaceous, such as ''P. speciosa'', ''P. rhomboidalis'', ''P. gradinarui'', etc. This genus is mainly a boreal taxon, being extensively reported in Europe, Iran, Afghanistan, China and North America, but it has been cited from Gondwanic occurrences as well, such as India, Argentina and Australia. ==References==
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