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Pterostichinae is a subfamily of ground beetles (family Carabidae). It belongs to the advanced harpaline assemblage, and if these are circumscribed ''sensu lato'' as a single subfamily, Pterostichinae are downranked to a tribe Pterostichini. However, as the former Pterostichitae supertribe of the Harpalinae as loosely circumscribed does seem to constitute a lineage rather distinct from ''Harpalus'', its core group is here considered to be the present subfamily and the Harpalinae are defined more narrowly. They are usually mid-sized and rather stout ground beetles. Coloration is typically dark and without conspicuous patterns, but often with a strong sheen like polished metal. They are widely distributed and inhabit a wide range of terrestrial habitats. Unlike the more basal ground beetles which only eat small animals, the Pterostichinae include a large proportion of omnivorous or even herbivorous taxa. ==Systematics== This group includes the following tribes and genera: Tribe Abacetini Chaudoir, 1872 * ''Abacetus'' Dejean, 1828 * ''Abacidus'' LeConte, 1873 * ''Aristopus'' Laferte-Senectere, 1853 * ''Celioinkosa'' Straneo, 1951 * ''Chlaeminus'' Motschulsky, 1865 * ''Colpodichius'' Straneo, 1952 * ''Cosmodiscus'' Sloane, 1907 * ''Cyrtomoscelis'' Chaudoir, 1874 * ''Distrigidius'' Jeannel, 1948 * ''Ecnomolaus'' Bates, 1892 * ''Haptoderidius'' Straneo, 1975 * ''Haptoderodes'' Straneo, 1986 * ''Holconotus'' Schmidt-Goebel, 1846 * ''Inkosa'' Peringuey, 1926 * ''Mateuellus'' Deuve, 1990 * ''Metabacetus'' Bates, 1892 * ''Metaxellus'' Straneo, 1960 * ''Metaxys'' Chaudoir, 1857 * ''Novillidius'' Straneo, 1941 * ''Oodinkosa'' Straneo, 1939 * ''Ophonichius'' Straneo, 1942 * ''Pediomorphus'' Chaudoir, 1878 * ''Pioprosopus'' Tschitscherine, 1899 * ''Pollicobius'' Vinson, 1939 * ''Prostalomus'' Basilewsky, 1950 * ''Pseudabacetus'' Burgeon, 1935 * ''Pterostillichus'' Straneo, 1949 * ''Rhagadillius'' Straneo, 1951 * ''Tiferonia'' Darlington, 1962 * ''Trachelocyphoides'' Alluaud, 1934 * ''Trachelocyphus'' Tschitscherine, 1900 Tribe Catapieseini Bates, 1882 * ''Catapiesis'' Solier, 1835 * ''Homalomorpha'' Brulle, 1835 Tribe Chaetodactylini Tschitscherine, 1903 * ''Chaetodactyla'' Tschitscherine, 1897 Tribe Cnemalobini Germain, 1911 * ''Cnemalobus'' Guerin-Meneville, 1838 Tribe Cratocerini Lacordaire, 1854 * ''Brachidius'' Chaudoir, 1852 * ''Cratocerus'' Dejean, 1829 * ''Oxyglychus'' Straneo, 1938 Tribe Drimostomatini Chaudoir, 1872 * ''Abacaelostus'' Straneo, 1952 * ''Andrewesinulus'' Straneo, 1938 * ''Apsidocnemus'' Alluaud, 1936 * ''Barylaus'' Liebherr, 1985 * ''Caecocaelus'' Straneo, 1949 * ''Caelostomus'' W.S. MacLeay, 1825 * ''Camptogenys'' Tschitscherine, 1899 * ''Capabatus'' Csiki, 1930 * ''Crenulostrigus'' Straneo, 1942 * ''Cyrtolaus'' Bates, 1882 * ''Dactyleurys'' Tschitscherine, 1899 * ''Dactylinius'' Straneo, 1941 * ''Diachipteryx'' Alluaud, 1926 * ''Diceromerus'' Chaudoir, 1872 * ''Dromistomus'' Jeannel, 1948 * ''Drymonaxus'' Straneo, 1941 * ''Eurystomis'' Chaudoir, 1878 * ''Feostoma'' Straneo, 1941 * ''Hemitelestus'' Alluaud, 1895 * ''Hoplizomenus'' Chaudoir, 1872 * ''Leleuporites'' Straneo, 1960 * ''Madapelmus'' Dajoz, 1985 * ''Monodryxus'' Straneo, 1942 * ''Pachycaecus'' Straneo, 1971 * ''Pachyroxochus'' Straneo, 1942 * ''Platyxythrius'' Straneo, 1942 * ''Stegazopteryx'' Will, 2004 * ''Stomonaxellus'' Tschitscherine, 1901 * ''Strigomerodes'' Straneo, 1939 * ''Strigomerus'' Chaudoir, 1872 * ''Trichillinus'' Straneo, 1938 Tribe Microcheilini Jeannel, 1948 * ''Microcheila'' Brulle, 1834 Tribe Morionini Brulle, 1835 * ''Buderes'' Murray, 1857 * ''Hyperectenus'' Alluaud, 1935 * ''Hyperion'' Castelnau, 1834 * ''Megamorio'' Chaudoir, 1880 * ''Morion'' Latreille, 1810 * ''Morionidius'' Chaudoir, 1880 * ''Moriosomus'' Motschulsky, 1864 * ''Platynodes'' Westwood, 1846 * ''Stereostoma'' Murray, 1857 Tribe Pterostichini Bonelli, 1810 (including Metiini) * ''Abacillius'' Straneo, 1949 * ''Abacillodes'' Straneo, 1988 * ''Abacoleptus'' Fauvel, 1903 * ''Abacomorphus'' Chaudoir, 1878 * ''Abacophrastus'' Will, 2011 * ''Abacops'' Tschitscherine, 1902 * ''Abaris'' Dejean, 1831 * ''Abax'' Bonelli, 1810 * ''Abropus'' Waterhouse, 1842 * ''Acanthoferonia'' Moore, 1965 * ''Adrimus'' Bates, 1871 * ''Allotriopus'' Bates, 1882 * ''Antarctiola'' Straneo, 1977 * ''Apsaustodon'' Tschitscherine, 1901 * ''Argutoridius'' Chaudoir, 1876 * ''Aristochroa'' Tschitscherine, 1898 * ''Aristochroodes'' Marcillac, 1993 * ''Aulacopodus'' Britton, 1940 * ''Basilewskya'' Straneo, 1948 * ''Blennidus'' Motschulsky, 1865 * ''Blenniventer'' Straneo, 1986 * ''Bothynoproctus'' Tschitscherine, 1900 * ''Camptoscelis'' Dejean, 1828 * ''Castelnaudia'' Tschitscherine, 1890 * ''Catadromus'' W.S. Macleay, 1825 * ''Cedrorum'' Borges & Serrano, 1993 * ''Cephalostichus'' Straneo, 1977 * ''Cerabilia'' Castelnau, 1867 * ''Chaetauchenium'' Tschitscherine * ''Chalcochrous'' Chaudoir, 1838 * ''Conchitella'' Moore, 1962 * ''Cophosomorpha'' Tschitscherine, 1890 * ''Cratoferonia'' Tschitscherine, 1902 * ''Cratogaster'' Blanchard, 1843 * ''Cyclotrachelus'' Chaudoir, 1838 * ''Cynthidia'' Chaudoir, 1873 * ''Darodilia'' Castelnau, 1867 * ''Eucamptognathus'' Chaudoir, 1837 * ''Euchroa'' Brulle, 1834 * ''Eudromus'' Klug, 1835 * ''Eumara'' Tschitscherine, 1901 * ''Euryabax'' Fauvel, 1903 * ''Euryaptus'' Bates, 1892 * ''Feroniola'' Tschitscherine, 1900 * ''Feronista'' Moore, 1965 * ''Gastrellarius'' Casey, 1918 * ''Gastrogmus'' Sloane, 1915 * ''Gourlayia'' Britton, 1964 * ''Haplobothynus'' Tschitscherine, 1901 * ''Haploferonia'' Darlington, 1962 * ''Harpostomus'' Chaudoir, 1856 * ''Henrotius'' Jeannel, 1953 * ''Holcaspis'' Chaudoir, 1865 * ''Homalonesiota'' Maindron, 1908 * ''Hybothecus'' Chaudoir, 1874 * ''Kuschelinus'' Straneo, 1963 * ''Leiradira'' Castelnau, 1867 * ''Lesticus'' Dejean, 1828 * ''Liopasa'' Tschitscherine, 1901 * ''Litarthrum'' Sloane, 1915 * ''Lobobrachus'' Sharp, 1885 * ''Lophoglossus'' LeConte, 1852 * ''Loxandrus'' LeConte, 1852 * ''Loxodactylus'' Chaudoir, 1865 * ''Loxogenius'' Sloane, 1907 * ''Marsyas'' Putzeys, 1845 * ''Mecynognathus'' W.J.MacLeay, 1873 * ''Megadromus'' Motschulsky, 1865 * ''Meropalpus'' Tschitscherine, 1900 * ''Metius'' Curtis, 1839 * ''Metoncidus'' Bates, 1871 * ''Microcephalus'' Dejean, 1828 * ''Mireius'' Straneo, 1984 * ''Molopidius'' Jeannel, 1942 * ''Molopinus'' Jeannel, 1948 * ''Molops'' Bonelli, 1810 * ''Myas'' Sturm, 1826 * ''Nebrioferonia'' Straneo, 1939 * ''Nelidus'' Chaudoir, 1878 * ''Neoferonia'' Britton, 1940 * ''Neotalus'' Will, 2002 * ''Nirmala'' Andrewes, 1930 * ''Notabax'' Moore, 1976 * ''Notolestus'' Sloane, 1894 * ''Notonomus'' Chaudoir, 1862 * ''Nurus'' Motschulsky, 1866 * ''Oberthueria'' Vuillet, 1911 * ''Ogmophora'' Tschitscherine, 1896 * ''Oribazus'' Chaudoir, 1874 * ''Orthomus'' Chaudoir, 1838 * ''Oscadytes'' Lagar Mascaro, 1975 * ''Oxycrepis'' Reiche, 1843 * ''Pachymorphus'' Chaudoir, 1838 * ''Pachythecus'' Chaudoir, 1874 * ''Paniestichus'' Will, 2011 * ''Paranurus'' Tschitscherine, 1901 * ''Pareuryaptus'' Dubault, Lassalle & Roux, 2008 * ''Parhypates'' Motschulsky, 1866 * ''Pedius'' Motschulsky, 1850 * ''Percolaus'' Bates, 1882 * ''Percus'' Bonelli, 1810 * ''Peyrieraselus'' Deuve, 1981 * ''Phaenaulax'' Tschitscherine, 1898 * ''Piesmus'' LeConte, 1852 * ''Platycaelus'' Blanchard, 1843 * ''Platysmodes'' Fauvel, 1903 * ''Plocamostethus'' Britton, 1940 * ''Poecilinus'' Jeannel, 1948 * ''Poecilus'' Bonelli, 1810 * ''Prosopogmus'' Chaudoir, 1865 * ''Psegmatopterus'' Chaudoir, 1878 * ''Pseudabarys'' Chaudoir, 1873 * ''Pseudoceneus'' Tschitscherine, 1891 * ''Pterostichus'' Bonelli, 1810 * ''Rhabdotus'' Chaudoir, 1865 * ''Rhytisternus'' Chaudoir, 1865 * ''Sarticus'' Motschulsky, 1865 * ''Secatophus'' Castelnau, 1867 * ''Setalidius'' Chaudoir, 1878 * ''Setalimorphus'' Sloane, 1895 * ''Setalis'' Castelnau, 1867 * ''Simodontus'' Chaudoir, 1843 * ''Speluncarius'' Reitter, 1886 * ''Speomolops'' Patrizi, 1955 * ''Sphodrosomus'' Perroud & Montrouzier, 1864 * ''Stereocerus'' Kirby, 1837 * ''Steropanus'' Fairmaire, 1888 * ''Sthenocranion'' Tschitscherine, 1899 * ''Stolonis'' Motschulsky, 1866 * ''Stomis'' Clairville, 1806 * ''Straneostichus'' Sciaky, 1994 * ''Styracoderus'' Chaudoir, 1874 * ''Taphoxomimus'' Straneo, 1975 * ''Tapinopterus'' Schaum, 1858 * ''Teratotarsa'' Tschitscherine, 1893 * ''Teropha'' Castelnau, 1867 * ''Trichosternus'' Chaudoir, 1865 * ''Trigonognatha'' Motschulsky, 1858 * ''Trigonotoma'' Dejean, 1828 * ''Trirammatus'' Chaudoir, 1838 * ''Tropidocerus'' Chaudoir, 1878 * ''Wahlbergiana'' Bousquet, 2002 * ''Xenion'' Tschitscherine, 1902 * ''Zariquieya'' Jeannel, 1924 * ''Zeodera'' Castelnau, 1867 * ''Zeopoecilus'' Sharp, 1886 Tribe Zabrini Bonelli, 1810 * ''Amara'' Bonelli, 1810 * ''Pseudamara'' Lindroth, 1968 * ''Zabrus'' Clairville, 1806 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Pterostichinae」の詳細全文を読む スポンサード リンク
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