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The scarab beetle subfamily Scarabaeinae consists of species collectively called true dung beetles. Most of the beetles of this subfamily feed exclusively on dung. Well-known members include the genera ''Scarabaeus'' and ''Sisyphus'', and ''Phanaeus vindex''. Scarabaeinae tribes, with some selected genera and species, are: * Canthonini * *Anachalcos Hope, 1837 * * ''Circellium'' Latreille, 1825 * * ''Dicranocara'' Frolov and Scholtz, 2003 * * ''Versicorpus'' Deschodt Davis and Scholtz, 2011 * * ''Deltochilum'' * * * ''Deltochilum valgum'' * Coprini * * ''Copris'' Geoffroy, 1762 * * ''Heliocopris'' * Dichotomiini * * ''Dichotomius'' * Eucraniini * Eurysterini * Gymnopleurini (sometimes included in Scarabaeini) * * ''Gymnopleurus'' Illiger, 1803 * Oniticellini * * ''Euoniticellus'' Janssens, 1953 * * ''Oniticellus'' Serville, 1825 * Onitini * * ''Bubas'' Mulsant, 1842 * * ''Cheironitis'' Lansberge, 1875 * * ''Onitis'' Fabricius, 1798 * Onthophagini * Phanaeini MacLeay, 1819 * * ''Oxysternon'' * * ''Phanaeus'' * * * ''Phanaeus changdiazi'' * * * ''Phanaeus vindex'' * Scarabaeini * * ''Scarabaeus'' * Sisyphini (sometimes included in Scarabaeini) * * ''Neosisyphus'' * * ''Sisyphus'' * Unknown tribe / ''Incertae sedis'' * * ''Anisocanthon'' * * ''Anomiopus'' Coprini, Oniticellini, Onitini, Onthophagini and Phanaeini are sometimes separated as subfamily Coprinae. File:Cheironitis scabrosus Fabricius 1776 female Syn. Chironitis scabrosus Fabricius, 1776 (7041225693).jpg|''Cheironitis scabrosus'' File:Chironitis indicus Lansberge, 1875 male (2872657334).jpg|''Chironitis indicus'' File:Copris elphenor Klug 1855 female (4011341553).jpg|''Copris elphenor'' female File:Copris elphenor Klug, 1855 male (4012106288).jpg|''Copris elphenor'' male ==External links== *(Scarabaeinae Research Network ) *(photos of species from each tribe ) *(African Dung Beetles Online ) 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Scarabaeinae」の詳細全文を読む スポンサード リンク
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