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'' Tetraxylopteris'' is a genus of extinct vascular plants of the Middle to Upper Devonian (around ). Fossils were first found in New York State, USA. A second species was later found in Venezuela. ==Description== Fossils of ''Tetraxylopteris'' have so far been discovered in two locations. ''T. schmidtii'' was named from the Catskill Clastic Wedge, New York State, United States of America, in rocks of Middle to Upper Devonian age (around ).〔 ''T. reposana'' was found in the Campo Chico Formation, north-west Venezuela, in beds believed to be of Frasnian age ().〔 The overall shape of ''Tetraxylopteris'' consisted of a complex system of branches. The main stem was 'pseudomonopodial', i.e. it divided dichotomously to produce side stems while the main stem maintained its identity. The main and side stems then bore three orders of branches (i.e. the first branches from the stems divided twice more). (There were possibly four orders in ''T. reposana''.) The branches were arranged in opposite pairs with successive pairs at right angles (decussate). In ''T. schmidtii'', the final order of branches bore appendages arranged oppositely and decussately; in ''T. reposana'', the appendages were opposite and decussate on the penultimate order of branches and spirally or helically arranged on the final order. The appendages branched dichotomously up to three times and were three-dimensional, although some appear flattened in the fossilized specimens.〔〔, pp. 489–481〕 They have been called "non-laminate proto-leaves" by Beerling and Fleming, reflecting the belief that such structures were precursors to true leaves, which evolved by first '' – flattening to produce a two-dimensional branched structure – and then 'webbing' – tissue growing out between the flattened branches. One consistent difference between the two species is that ''T. schmidtii'' had branches which very distinctly narrowed along their length whereas only the third order branches of ''T. reposana'' showed any tapering. Another is that ''T. reposana'' had swellings at the bases of first and second order branches.〔 In both species, the primary xylem strand was central to stems, branches and appendages. In the main stems and branches it was X-shaped in cross-section, corresponding to the four rows of branches. In the final appendages it became circular in cross-section. The xylem development was 'mesarch', i.e. the first maturing protoxylem had later maturing metaxylem on either side. Protoxylem occurred both at the tips of the lobes of the xylem strand and in the centre. The general anatomy of the woody stem resembles that of seed plants.〔〔 The spore-forming organs or sporangia of ''Tetraxylopteris'' were born on a very complex 'fertile branching system'. Firstly the main axis of the system branched twice dichotomously. Then each of the four resulting branches was three times pinnate. Each ultimate unit had an elongated sporangium at its end which split longitudinally to release the spores which were trilete, ranging from around 70 to 170 µm in diameter.〔 The complex three-dimensional branching pattern imples that both species would have been open bushy plants. The exact height cannot be determined from the fossils, which consist of broken-off portions. The longest known section of ''T. schmidtii'' is 50 cm – rather less for ''T. reposana'', which may nevertheless have been the taller species. An overall height of a few metres has been proposed. Hammond and Berry suggest that ''T. reposana'' may have grown in dense thickets so that plants supported one another.〔 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Tetraxylopteris」の詳細全文を読む スポンサード リンク
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