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Tomarus
''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.〔Ratcliffe, B. C. and R. D. Cave. (2010). (The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of ''Tomarus adoceteus'', new species (Pentodontini) and ''Caymania nitidissima'', new genus and species (Phileurini). ) ''Insecta Mundi'' 139, 1-15.〕 The adult beetles are nocturnal and attracted to lights; the larvae generally remain in the soil, often feeding on plant roots.〔 In order to identify species, the parameres of the male must be pulled out of the abdomen and examined. This process should be done carefully, because the parameres are quite fragile, "almost parchment-like".〔 There are about 27 species in the genus.〔 Species include:〔〔Ratcliffe, B. C. (2003). (The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). ) ''Bulletin of the University of Nebraska State Museum'' 16, 1-506.〕 *''Tomarus adoceteus'' *''Tomarus bituberculatus'' *''Tomarus cicatricosus'' *''Tomarus cuniculus'' *''Tomarus ebenus'' *''Tomarus fossor'' *''Tomarus gyas'' *''Tomarus laevicollis'' *''Tomarus maternus'' *''Tomarus nasutus'' *''Tomarus sallaei'' *''Tomarus similis'' *''Tomarus subtropicus'' – sugarcane grub〔Buss, E. A. (Sugarcane Grub, ''Tomarus subtropicus'' Blatchley (Insecta: Coleoptera: Scarabaeidae). ) EENY-318 (IN593). Entomology and Nematology. Florida Cooperative Extension Service. University of Florida IFAS. Published 2003, revised 2006.〕 ==References==
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