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Tupuxuara : ウィキペディア英語版
Tupuxuara

''Tupuxuara'' is a genus of large, crested, toothless pterodactyloid pterosaur.
==Discovery==


The genus was named and described by Alexander Kellner and Diógenes de Almeida Campos in 1988.〔Kellner, A.W.A., and Campos, D.A. (1988). "Sobre un novo pterossauro com crista sagital da Bacia do Araripe, Cretaceo Inferior do Nordeste do Brasil. (Pterosauria, Tupuxuara, Cretaceo, Brasil)." ''Anais de Academia Brasileira de Ciências'', 60: 459–469. (Portuguese )〕 The type species is ''Tupuxuara longicristatus''. The genus name refers to a familiar spirit from the mythology of the Tupi. The specific name means "long-crested" in Latin.
The holotype, MN 6591-V, was found in the Cretaceous Santana Formation of Brazil. It consists of a snout and some partial wing bones. Mature individuals of ''T. longicristatus'' had a back-swept crest arising from the snout. Much more fossil material has later been found, showing considerable variation in morphology. Some researchers explain this as intra-specific variability, being caused by a difference in age or sex. Others, however, assume there are different species present.
In 1994 a second species was named by Kellner: ''Tupuxuara leonardii''. The specific name honours Giuseppe Leonardi. The holotype is MN 6592-V, a fragmentary skull with a more rounded crest. Other such material has been referred to ''T. leonardii''. The largest skulls have a length of 130 centimetres indicating a wingspan of 5.5 metres (18 ft).
In 2009 a third species was named, by Mark Paul Witton: ''Tupuxuara deliradamus''. The holotype is SMNK PAL 6410, a skull. Another skull is the paratype: KPMNH DL 84. The specific name is derived from Latin ''delirus'', "insane" or "crazy", and ''adamas'', "invincible" but also the word from which "diamond" is derived. The species has a distinctive diamond-shaped skull opening and low eye sockets. The name is a tribute to the song "Shine On You Crazy Diamond" by Pink Floyd, one of Witton's favourite bands.〔Witton, M.P. (2009). "A new species of ''Tupuxuara'' (Thalassodromidae, Azhdarchoidea) from the Lower Cretaceous Santana Formation of Brazil, with a note on the nomenclature of Thalassodromidae", ''Cretaceous Research'' 30'(5): 1293-1300〕

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