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Urticales is a botanical name for what used to be an order of flowering plants. Before molecular phylogenetics became an important part of plant taxonomy, Urticales was recognized in many, perhaps even most, systems of plant classification, with some variations in circumscription. Among these is the Cronquist system (1981), which placed the order in the subclass Hamamelidae , as comprising: * Barbeyaceae * Cannabaceae * Cecropiaceae * Moraceae * Ulmaceae * Urticaceae In the APG III system (2009), the plants belonging to this order, along with four other families, constitute the order Rosales. Cecropiaceae is no longer recognized as separate from Urticaceae. The families Ulmaceae, Cannabaceae, Moraceae, and Urticaceae form a clade that has strong statistical support in phylogenetic analyses of DNA sequences.〔Shu-dong Zhang, Douglas E. Soltis, Yang Yang, De-zhu Li, and Ting-shuang Yi. "Multi-gene analysis provides a well-supported phylogeny of Rosales". ''Molecular Phylogenetics and Evolution'' 60(1):21-28.〕 This clade has been informally called the urticalean rosids.〔Kenneth J. Sytsma, Jeffery Morawetz, J. Chris Pires, Molly Nepokroeff, Elena Conti, Michelle Zjhra, Jocelyn C. Hall, and Mark W. Chase. (2002). Urticalean rosids: Circumscription, rosid ancestry, and phylogenetics based on ''rbcL'', ''trnL-F'', and ''ndhF'' sequences. ''American Journal of Botany'' 89(9): 1531-1546. (PDF fulltext )〕 Urticaleean rosids refers to the relationships amongst several families of angiosperms, and now includes more than 2,500 species. The families are: * Celtidaceae * Cannabaceae * Cecropiaceae * Moraceae * Ulmaceae * Urticaceae〔Qiu Y.-L. M. W. Chase S. B. Hoot E. Conti P. R. Crane K. J. Sytsma C. R. Parks 1998. Phylogenetics of the Hamamelidae and their allies: parsimony analyses of nucleotide sequences of the plastid gene rbcL. International Journal of Plant Sciences 159: 891-905.〕 The relationships within the "urticalean lineage" are now considered to be within the Rosales.〔APG (Angiosperm Phylogeny Group). 1998. An ordinal classification for the families of flowering plants. Annals of the Missouri Botanical Garden 85: 531-553.〕 The Moraceae and Urticaceae account for approximately 90% of the diversity within the clade.〔Sytsma, Kenneth J., Jeffery Morawetz, J. Chris Pires, Molly Nepokroeff, Elena Conti, Michelle Zjra, Jocelyn C. Hall, and Mark W. Chase. 2001. Urticalean Rosids: Circumscription, Rosid Ancestry, and Phylogenetics Based on RbcL, TrnL-F, and NdhF Sequences. American Journal of Botany〕 Analysis of DNA samples in ''rbcL'', ''trnL-F'', and ''ndhF'' plastid regions suggests that Urticalean rosids are derived out of a lineage including Barbeyaceae, Dirachmaceae, Elaegnaceae, and Rhamnaceae, with Rosaceae less closely related.〔 The morphological and molecular characters which define "urticalean rosins" are: * A dense gray tomentum comprising curly unicellular trichomes on abaxial leaf surfaces 〔Tobe H. and T. Takaso 1996. Trichome micromorphology in Celtidaceae and Ulmaceae (Urticales). Acta Phytotaxonomica et Geobotanica 47: 153–168.〕 * A reduced stamen count to one whorl or less〔Judd W. S. C. S. Campbell E. A. Kellogg P. F. Stevens 1999. Plant systematics: a phylogenetic approach. Sinauer Associates, Sunderland, Massachusetts, USA.〕〔Stevens P. F. 2001. Angiosperm phylogeny website, v. 2, August 2001. http://www.mobot.org/MOBOT/research/APweb/.〕 * Presence of inconspicuous flowers with up to five stamens〔〔 * Presence of two carpels〔〔 * A unilocular ovary with single apical ovule〔〔 * Urticoid teeth upon leaves 〔〔 * Developed prophyllar buds between paired inflorescences〔〔 Urticalean families span a wide range of morphological features—deciduous or evergreen trees, vines, shrubs, annuals and some succulents. Leaves also vary though they tend to share brochidodromous or palmately pinnate venation, often associated with lobing or compounding in the leaf blade. Among Moraceae, Urticacae and Cecropiaceae, mucilage cells and latex production is common. However, Cannabaceae, Ulmaceae, and Celitdaceae do not produce this material despite the presence of laticifers. == References == 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Urticales」の詳細全文を読む スポンサード リンク
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