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frond
A frond is a large, divided leaf.〔Raven, Evert Eichhorn,2004. The Biology of Plants, 7th Ed. W.H. Freeman and Company, New York, NY.〕 In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds〔Gifford and Foster, 1989. Morphology and Evolution of Vascular Plants, 3rd Ed. W.H. Freeman and Company, New York, NY.〕 and some botanists restrict the term to this group.〔Judd, Campbell, Kellogg, Stevens, Donoghue, 2007. Plant Systematics: A Phylogenetic Approach, 3rd Ed. Sinauer, Sunderland, MA.〕 Other botanists allow the term frond to also apply to the large leaves of cycads and palms (Arecaceae).〔Jones, 1993. Cycads of the World. Smithsonian Institution Press,USA.〕〔Allaby, 1992. The Concise Oxford Dictionary of Botany. Oxford University Press, Oxford, UK.〕 "Frond" is commonly used to identify a large, compound leaf, but if the term is used botanically to refer to the leaves of ferns, it may be applied to smaller and undivided leaves.
Fronds have particular terms describing their components. Like all leaves, fronds usually have a stalk connecting them to the main stem. In botany, this leaf stalk is generally called a petiole, but in regard to fronds specifically it is called a stipe, and it supports a flattened blade (which may be called a lamina), and the continuation of the stipe into this portion is called the rachis. The blades may be simple (undivided), pinnatifid (deeply incised, but not truly compound), pinnate (compound with the leaflets arranged along a rachis to resemble a feather), or further compound (subdivided). If compound, a frond may be compound once, twice, or more.
==Pinnate fronds==
If a frond is pinnate, each leafy segment of the blade is called a pinna (plural pinnae), the stalk bearing the pinna a petiolule, and the main vein or mid-rib of the pinna a costa (plural costae).〔Walters, Keil, 1996. Vascular Plant Taxonomy, 4th Ed. Kendall Hunt Publishing Co. Dubuque, IA.〕 If a frond is divided into pinnae, the frond is called once pinnate. In some fronds the pinna are further divided into segments, creating a bipinnate frond. The segments into which each pinna are divided are called pinnules. Rarely, a frond may even be tripinnate, in which case the pinnule divisions are known as ultimate segments. Pinnae may be arranged along the rachis either directly opposite one another or alternating up the stem. The arrangement may change from the base of a blade to the tip, as in the example of ''Blechnum'' shown below (from base to tip: pinnae opposite to alternate, and pinnatisect to pinnatifid).
Some fronds are not pinnately compound (or simple), but may be palmate or bifurcate. Some ferns, like members of the group Ophioglossales have a unique arrangement.
抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』
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