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hongshanornis : ウィキペディア英語版
hongshanornis

''Hongshanornis'' is a genus of ornithuromorph birds known from early Cretaceous lake deposits of the Yixian Formation, Inner Mongolia, China. The holotype specimen, recovered in 2005, is currently held by the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing. It was found in the Jianshangou fossil beds, dated to 124.6 million years ago. Three additional specimens have been reported, though only one of those has been definitively identified as belonging to ''Hongshanornis''. This latter specimen was found in the Dawangzhangzi fossil beds, which are about 122 million years old.〔
''Hongshanornis'' is a member of the group Hongshanornithidae, to which it lent its name. It is closely related to ''Longicrusavis'', which existed alongside ''Hongshanornis'' in the Dawangzhangzi ecosystem, and is very similar to the later ''Parahongshanornis'' from the Jiufotang Formation.〔
==Description==
''Hongshanornis longicresta'' was a small species, especially compared to other early ornithuromorphs (birds with a modern tail anatomy), about the size of a thrush, and adult specimens are estimated to have weighed about in life, with a wingspan of about .〔
The skull in all known specimens is poorly preserved, but in general appears to have had a narrow snout compared to the closely related ''Longicrusavis''.〔 The teeth were very small and are poorly preserved in all known specimens. At first, this led scientists to conclude that the teeth were absent in both the upper and lower jaws, probably replaced with a beak. However, later study of the type specimen showed that tooth sockets were present in the preserved parts of the jaw, and comparison with related forms showed that it did have teeth. The discovery of a more well preserved specimen confirmed the presence of teeth in at least the maxilla (middle of the upper jaw) and dentary (lower jaw). The snout appears to have lacked feathers, though whether it was only covered in skin or some parts bore a beak is currently unknown. The describers of ''Hongshanornis'' noted a unique bone in the end of the lower jaw which they equated with the beak-bearing "predentary bone" similar to the one found in ornithischian dinosaurs. Such bones have also been found in more advanced birds like ''Hesperornis''.〔 (2005): Discovery of an ornithurine bird and its implication for Early Cretaceous avian radiation. ''PNAS'' 102(52): 18998-19002. PMID 16344487 - Fulltext: (HTML ), (PDF ), (Supporting Information )〕 However, other scientists have pointed out that the "predentary" (or, technically, ''mandibular symphysial ossification'') of ''Hongshanornis'' lacks the characteristic pits and grooves associated with the beak in early beaked birds like ''Archaeorhynchus'', and that the presence or absence of any beak is unclear.〔〔
The original describers of ''Hongshanornis'' noted the apparent presence of a large feathered crest on the head,〔 though the feather traces are extremely poor quality and it may be an artifact of preservation.〔 A second specimen from different fossil beds did not have long feathers on the head, but rather showed feathers which became shorter closer to the snout.〔
The wings were long and broad, but tapered and pointed at the tips. The first few primary flight feathers were much shorter than the rest. The total wing area was 0.016 square meters. The wing digits bore small claws, with the claw on the first finger (the alular digit) much larger than that of the second finger (major digit).〔 ''Hongshanornis'' is characterized by very long legs, with the lower leg (''tibiotarsus'') more than one and a half times longer than the upper leg (''femur''). The toes were long and thin, with small claws. The first toe (''hallux'') was very small, and the second toe was much shorter than the fourth. The third toe was longest.〔
The bones of the tail have not been preserved in any known specimen, but comparison with related species suggest it was short with a small, plowshare-shaped pygostyle bone, to which the tail feathers anchored. A mass of muscles, called the rectorial bulb, probably was present to control the fanning of the trail feathers. The tail feathers were long, with symmetrical vanes. There were at least 10 tail feathers (''rectrices''), more than in other primitive fan-tailed birds. The overall shape of the tail was rounded and wedge-shaped when viewed from above; due to the symmetrical feathers, the sides of the tail were probably not held fully fanned into the wind but swept back, forming a partially fanned wedge as in some modern birds like accipiters, flycatchers and sunbitterns.〔

抄文引用元・出典: フリー百科事典『 ウィキペディア(Wikipedia)
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