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raphide
Raphides are needle-shaped crystals of calcium oxalate as the monohydrate or calcium carbonate as aragonite, found in more than 200 families of plants.
Both ends are needle-like, but raphides tend to be blunt at one end and sharp at the other.
== Calcium oxalate in plants ==
Many plants accumulate calcium oxalate crystals in response to surplus calcium, which is found throughout the natural environment. The crystals are produced in an intriguing variety of shapes. The crystal morphology depends on the taxonomic group of the plant. In one study of over 100 species, it was found that calcium oxalate accounted for 6.3% of plant dry weight. Crystal morphology and the distribution of raphides (in roots or leaves or tubers etc.) is similar in same taxa but different in others leaving possible opportunities for plant key characteristics and systematic identification; mucilage in raphide containing cells makes light microscopy difficult, though. Little is known about the mechanisms of sequestration or indeed the reason for accumulation of raphides but it is most likely as a defense mechanism against herbivory. It has also been suggested that in some cases raphides may help form plant skeletal structure. Raphides typically occur in parenchyma cells in aerial organs especially the leaves, and are generally confined to the mesophyl. As the leaf area increases, so does the number of raphides, the process starting in even young leaves. The first indications that the cell will contain crystals is shown when the cells enlarge with a larger nucleus.
Raphides are found in specialized plant cells or crystal chambers called idioblasts.〔 Also, 〕 Electron micrographs have shown that raphide needle crystals are normally four sided or H-shaped (with a groove down both sides) or with a hexagonal cross section and some are barbed. Wattendorf (1976) suggested that all circular sectioned raphides, as visible in a light microscope, are probably hexagonal in cross section Microscopy using polarized light shows bright opalescence with raphides. Plants like Tradescantia pallida also accumulate calcium oxalate crystals in response to heavy metals stress.
抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』
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